history of email

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Next, the CPYNET code was incorporated into SNDMSG. It remained to provide a way to distinguish local mail from network mail. I chose to append an at sign and the host name to the user’s (login) name. I am frequently asked why I chose the at sign, but the at sign just makes sense. The purpose of the at sign (in English) was to indicate a unit price (for example, 10 items @ $1.95). I used the at sign to indicate that the user was “at” some other host rather than being local.
The first message was sent between two machines that were literally side by side. The only physical connection they had (aside from the floor they sat on) was through the ARPANET. I sent a number of test messages to myself from one machine to the other. The test messages were entirely forgettable and I have, therefore, forgotten them. Most likely the first message was QUERTYIOP or something similar. When I was satisfied that the program seemed to work, I sent a message to the rest of my group explaining how to send messages over the network. The first use of network email announced its own existence.
These first messages were sent in late 1971. The next release of TENEX went out in early 1972 and included the version of SNDMSG with network mail capabilities. The CPYNET protocol was soon replaced with a real file transfer protocol having specific mail handling features. Later, a number of more general mail protocols were developed.
The First Network Email (openmap.bbn.com)
The first use of network email announced its own existence. Ray Tomlinson, The First Network Email

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History of email (Wikipedia) The history of email entails an evolving set of technologies and standards that culminated in the email systems in use today.